:root {
  --ink: #142820;
  --forest: #173a2d;
  --forest-deep: #0c261d;
  --moss: #59755d;
  --leaf: #a9bf91;
  --acid: #f1cf3e;
  --acid-soft: #f5df77;
  --cream: #f5f1e7;
  --paper: #fbfaf6;
  --white: #fff;
  --line: rgba(20, 40, 32, 0.14);
  --shadow: 0 24px 60px rgba(16, 44, 33, 0.12);
  --serif: "Iowan Old Style", "Palatino Linotype", "Book Antiqua", Georgia, serif;
  --sans: "Avenir Next", Avenir, "Segoe UI", Helvetica, Arial, sans-serif;
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  color: var(--ink);
  background: var(--paper);
  font-family: var(--sans);
  font-size: 16px;
  line-height: 1.65;
  -webkit-font-smoothing: antialiased;
}

body,
a,
button,
summary {
  -webkit-tap-highlight-color: transparent;
}

img {
  display: block;
  max-width: 100%;
}

a {
  color: inherit;
  text-decoration: none;
}

h1,
h2,
h3,
p {
  margin-top: 0;
}

h1,
h2 {
  font-family: var(--serif);
  font-weight: 500;
  letter-spacing: -0.045em;
  line-height: 0.98;
}

h2 {
  margin-bottom: 24px;
  font-size: clamp(2.45rem, 5vw, 4.8rem);
}

h3 {
  line-height: 1.25;
}

.shell {
  width: min(1180px, calc(100% - 48px));
  margin-inline: auto;
}

.section {
  padding: 120px 0;
}

.skip-link {
  position: fixed;
  z-index: 200;
  top: 12px;
  left: 12px;
  padding: 10px 16px;
  color: var(--forest-deep);
  background: var(--acid);
  border-radius: 5px;
  transform: translateY(-160%);
}

.skip-link:focus {
  transform: none;
}

.site-header {
  position: absolute;
  z-index: 50;
  top: 0;
  right: 0;
  left: 0;
  color: white;
  border-bottom: 1px solid rgba(255, 255, 255, 0.2);
}

.header-inner {
  display: flex;
  align-items: center;
  min-height: 92px;
  gap: 34px;
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 11px;
  font-size: 1.35rem;
  letter-spacing: -0.04em;
  line-height: 1;
  white-space: nowrap;
}

.brand strong {
  color: var(--acid);
  font-weight: 800;
}

.brand small {
  font-size: 0.72em;
  opacity: 0.62;
}

.brand-mark {
  position: relative;
  display: inline-block;
  width: 32px;
  height: 32px;
  border: 1px solid rgba(255, 255, 255, 0.62);
  border-radius: 48% 52% 61% 39% / 42% 45% 55% 58%;
  transform: rotate(-12deg);
}

.brand-mark i {
  position: absolute;
  width: 6px;
  height: 6px;
  background: var(--acid);
  border-radius: 50%;
}

.brand-mark i:nth-child(1) { top: 6px; left: 8px; }
.brand-mark i:nth-child(2) { top: 15px; right: 6px; }
.brand-mark i:nth-child(3) { bottom: 5px; left: 7px; }

.desktop-nav {
  display: flex;
  align-items: center;
  justify-content: center;
  gap: 30px;
  margin-left: auto;
  font-size: 0.88rem;
}

.desktop-nav a {
  position: relative;
  padding: 10px 0;
  opacity: 0.86;
}

.desktop-nav a::after {
  position: absolute;
  right: 0;
  bottom: 2px;
  left: 0;
  height: 1px;
  background: var(--acid);
  content: "";
  transform: scaleX(0);
  transform-origin: right;
  transition: transform 180ms ease;
}

.desktop-nav a:hover::after,
.desktop-nav a:focus-visible::after {
  transform: scaleX(1);
  transform-origin: left;
}

.mobile-menu {
  display: none;
  margin-left: auto;
}

.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 54px;
  padding: 0 23px;
  gap: 14px;
  color: var(--forest-deep);
  background: var(--acid);
  border: 1px solid var(--acid);
  border-radius: 999px;
  font-size: 0.9rem;
  font-weight: 700;
  letter-spacing: 0.01em;
  transition: transform 180ms ease, background-color 180ms ease, border-color 180ms ease;
}

.button:hover,
.button:focus-visible {
  background: var(--acid-soft);
  border-color: var(--acid-soft);
  transform: translateY(-2px);
}

.button-small {
  min-height: 42px;
  padding-inline: 19px;
  font-size: 0.82rem;
}

.text-link {
  display: inline-flex;
  align-items: center;
  width: fit-content;
  gap: 10px;
  padding-bottom: 4px;
  border-bottom: 1px solid currentColor;
  font-size: 0.86rem;
  font-weight: 700;
}

.text-link span {
  transition: transform 180ms ease;
}

.text-link:hover span,
.text-link:focus-visible span {
  transform: translateX(4px);
}

.light-link {
  color: white;
  border-color: rgba(255, 255, 255, 0.52);
}

.eyebrow {
  margin-bottom: 18px;
  color: var(--moss);
  font-size: 0.72rem;
  font-weight: 800;
  letter-spacing: 0.18em;
  text-transform: uppercase;
}

.eyebrow.light {
  color: var(--acid-soft);
}

.hero {
  position: relative;
  display: grid;
  align-items: center;
  min-height: min(920px, 100vh);
  overflow: hidden;
  color: white;
  background: var(--forest-deep);
}

.hero-image,
.hero-shade {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
}

.hero-image {
  object-fit: cover;
  object-position: center;
}

.hero-shade {
  background:
    linear-gradient(90deg, rgba(6, 25, 18, 0.96) 0%, rgba(6, 25, 18, 0.82) 28%, rgba(6, 25, 18, 0.2) 64%, rgba(6, 25, 18, 0.03) 100%),
    linear-gradient(0deg, rgba(7, 25, 19, 0.56) 0%, transparent 36%);
}

.hero-inner {
  position: relative;
  z-index: 2;
  padding-top: 110px;
}

.hero-copy {
  width: min(690px, 66%);
}

.hero h1 {
  max-width: 680px;
  margin-bottom: 28px;
  font-size: clamp(3.8rem, 7.2vw, 7rem);
  text-wrap: balance;
}

.hero-lead {
  max-width: 590px;
  margin-bottom: 34px;
  color: rgba(255, 255, 255, 0.8);
  font-size: clamp(1.05rem, 1.8vw, 1.24rem);
  line-height: 1.6;
}

.button-row {
  display: flex;
  align-items: center;
  flex-wrap: wrap;
  gap: 26px;
}

.hero-note {
  position: absolute;
  z-index: 3;
  right: 0;
  bottom: 0;
  display: flex;
  align-items: center;
  max-width: 405px;
  min-height: 82px;
  padding: 18px 46px 18px 30px;
  gap: 13px;
  color: var(--ink);
  background: var(--cream);
  font-size: 0.82rem;
  line-height: 1.45;
}

.pulse-dot {
  width: 10px;
  height: 10px;
  flex: 0 0 auto;
  background: var(--acid);
  border: 3px solid rgba(23, 58, 45, 0.12);
  border-radius: 50%;
  box-sizing: content-box;
}

.intro {
  background: var(--paper);
}

.section-heading {
  margin-bottom: 62px;
}

.split-heading {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(280px, 0.65fr);
  align-items: end;
  gap: 90px;
}

.split-heading h2 {
  max-width: 760px;
  margin-bottom: 0;
}

.heading-copy {
  margin-bottom: 10px;
  color: #577065;
  font-size: 1.04rem;
}

.fact-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  border-top: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
}

.fact-card {
  position: relative;
  min-height: 430px;
  padding: 42px 42px 38px;
  border-right: 1px solid var(--line);
  transition: background-color 200ms ease, transform 200ms ease;
}

.fact-card:last-child {
  border-right: 0;
}

.fact-card:hover {
  background: white;
  transform: translateY(-6px);
}

.fact-card.featured {
  background: #f2efdf;
}

.card-number {
  color: #7c8b83;
  font-size: 0.68rem;
  font-weight: 800;
  letter-spacing: 0.16em;
}

.fact-card h3 {
  max-width: 260px;
  margin: 34px 0 15px;
  font-family: var(--serif);
  font-size: 1.8rem;
  font-weight: 500;
  letter-spacing: -0.035em;
}

.fact-card p {
  min-height: 84px;
  color: #5c7067;
  font-size: 0.93rem;
}

.fact-card .text-link {
  position: absolute;
  bottom: 40px;
}

.fact-symbol {
  position: absolute;
  top: 34px;
  right: 34px;
  width: 72px;
  height: 72px;
  opacity: 0.72;
}

.fact-symbol i {
  position: absolute;
  display: block;
}

.spores i {
  width: 16px;
  height: 16px;
  border: 1.5px solid var(--forest);
  border-radius: 50%;
}

.spores i::before,
.spores i::after {
  position: absolute;
  top: 6px;
  left: -5px;
  width: 4px;
  height: 4px;
  background: var(--forest);
  border-radius: 50%;
  content: "";
}

.spores i::after { right: -5px; left: auto; }
.spores i:nth-child(1) { top: 5px; left: 22px; }
.spores i:nth-child(2) { top: 29px; left: 7px; transform: scale(0.72); }
.spores i:nth-child(3) { top: 41px; left: 38px; transform: scale(1.2); }
.spores i:nth-child(4) { top: 15px; right: 4px; transform: scale(0.55); }

.route::before {
  position: absolute;
  top: 12px;
  left: 15px;
  width: 40px;
  height: 48px;
  border: 1.5px dashed var(--forest);
  border-top: 0;
  border-left-color: transparent;
  border-radius: 50%;
  content: "";
  transform: rotate(42deg);
}

.route i {
  width: 9px;
  height: 9px;
  background: var(--forest);
  border: 4px solid #e7e2ce;
  border-radius: 50%;
  box-sizing: content-box;
}

.route i:nth-child(1) { top: 7px; left: 7px; }
.route i:nth-child(2) { top: 27px; right: 3px; }
.route i:nth-child(3) { bottom: 2px; left: 22px; }

.shield i {
  top: 6px;
  left: 10px;
  width: 50px;
  height: 58px;
  border: 1.5px solid var(--forest);
  border-radius: 25px 25px 32px 32px / 20px 20px 42px 42px;
}

.shield i::before,
.shield i::after {
  position: absolute;
  background: var(--forest);
  content: "";
}

.shield i::before { top: 28px; left: 14px; width: 20px; height: 1px; transform: rotate(-45deg); }
.shield i::after { top: 32px; left: 11px; width: 9px; height: 1px; transform: rotate(45deg); }

.dark-section {
  color: white;
  background: var(--forest-deep);
}

.disease-grid {
  display: grid;
  grid-template-columns: minmax(0, 0.8fr) minmax(480px, 1.2fr);
  align-items: center;
  gap: 90px;
}

.disease-copy h2 {
  font-size: clamp(2.8rem, 5.4vw, 5.2rem);
}

.disease-copy > p:not(.eyebrow) {
  margin-bottom: 30px;
  color: rgba(255, 255, 255, 0.7);
}

.check-list {
  margin: 0 0 38px;
  padding: 0;
  list-style: none;
}

.check-list li {
  position: relative;
  padding: 13px 0 13px 34px;
  border-bottom: 1px solid rgba(255, 255, 255, 0.12);
  font-size: 0.9rem;
}

.check-list li::before {
  position: absolute;
  top: 18px;
  left: 4px;
  width: 13px;
  height: 7px;
  border-bottom: 1.5px solid var(--acid);
  border-left: 1.5px solid var(--acid);
  content: "";
  transform: rotate(-45deg);
}

.button-ghost {
  color: white;
  background: transparent;
  border-color: rgba(255, 255, 255, 0.45);
}

.button-ghost:hover,
.button-ghost:focus-visible {
  color: var(--forest-deep);
  background: white;
  border-color: white;
}

.disease-visual {
  position: relative;
  min-height: 620px;
}

.photo-card {
  position: absolute;
  z-index: 2;
  margin: 0;
  overflow: hidden;
  background: white;
  box-shadow: 0 30px 80px rgba(0, 0, 0, 0.36);
}

.photo-card img {
  width: 100%;
  height: 100%;
  object-fit: cover;
}

.photo-card figcaption {
  position: absolute;
  right: 0;
  bottom: 0;
  padding: 10px 14px;
  color: var(--ink);
  background: var(--acid);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.05em;
}

.photo-card-main {
  top: 0;
  right: 0;
  width: 80%;
  height: 460px;
}

.photo-card-main img {
  filter: contrast(1.08);
}

.photo-card-small {
  bottom: 0;
  left: 0;
  width: 56%;
  height: 285px;
  border: 10px solid var(--forest-deep);
}

.orbit {
  position: absolute;
  z-index: 1;
  display: block;
  border: 1px solid rgba(241, 207, 62, 0.32);
  border-radius: 50%;
}

.orbit-one { top: -35px; left: 10px; width: 410px; height: 410px; }
.orbit-two { right: -80px; bottom: -55px; width: 330px; height: 330px; }

.actions {
  background: var(--paper);
}

.centered-heading {
  max-width: 760px;
  margin-right: auto;
  margin-left: auto;
  text-align: center;
}

.centered-heading h2 {
  font-size: clamp(2.6rem, 5vw, 4.6rem);
}

.centered-heading > p:last-child {
  max-width: 630px;
  margin: 0 auto;
  color: #62756c;
}

.action-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 18px;
}

.action-card {
  position: relative;
  display: flex;
  flex-direction: column;
  justify-content: space-between;
  min-height: 480px;
  padding: 34px;
  overflow: hidden;
  transition: transform 200ms ease, box-shadow 200ms ease;
}

.action-card:hover,
.action-card:focus-visible {
  box-shadow: var(--shadow);
  transform: translateY(-8px);
}

.action-yellow { background: var(--acid); }
.action-cream { background: #e5ead7; }
.action-photo { color: white; background: var(--forest); }

.action-photo > img,
.photo-overlay {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
}

.action-photo > img {
  object-fit: cover;
}

.photo-overlay {
  background: linear-gradient(180deg, rgba(8, 30, 22, 0.08), rgba(8, 30, 22, 0.88));
}

.action-card > *:not(img):not(.photo-overlay) {
  position: relative;
  z-index: 2;
}

.action-index {
  font-size: 0.7rem;
  font-weight: 800;
  letter-spacing: 0.16em;
}

.mini-label {
  margin-bottom: 12px;
  font-size: 0.67rem;
  font-weight: 800;
  letter-spacing: 0.17em;
  text-transform: uppercase;
  opacity: 0.7;
}

.action-card h3 {
  max-width: 300px;
  margin-bottom: 13px;
  font-family: var(--serif);
  font-size: 2.1rem;
  font-weight: 500;
  letter-spacing: -0.04em;
}

.action-card p:last-child {
  margin-bottom: 50px;
  font-size: 0.92rem;
  opacity: 0.77;
}

.round-arrow {
  position: absolute !important;
  right: 30px;
  bottom: 28px;
  display: grid;
  width: 44px;
  height: 44px;
  place-items: center;
  border: 1px solid currentColor;
  border-radius: 50%;
  font-size: 1.25rem;
}

.captivity {
  background: #e7edde;
}

.captivity-grid {
  display: grid;
  grid-template-columns: minmax(0, 1.15fr) minmax(360px, 0.85fr);
  align-items: center;
  gap: 100px;
}

.captivity-media {
  position: relative;
  min-height: 660px;
}

.captivity-media > img {
  width: 92%;
  height: 620px;
  object-fit: cover;
  filter: saturate(0.76) contrast(0.98);
}

.image-label {
  position: absolute;
  top: 0;
  left: 0;
  padding: 12px 18px;
  color: var(--forest-deep);
  background: var(--acid);
  font-size: 0.7rem;
  font-weight: 800;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.drawn-ripples {
  position: absolute;
  right: -16px;
  bottom: -8px;
  width: 210px;
  height: 170px;
}

.drawn-ripples i {
  position: absolute;
  right: 0;
  bottom: 0;
  display: block;
  border: 1px solid rgba(20, 40, 32, 0.38);
  border-radius: 50%;
  transform: rotate(-8deg);
}

.drawn-ripples i:nth-child(1) { width: 210px; height: 90px; }
.drawn-ripples i:nth-child(2) { right: 25px; bottom: 32px; width: 150px; height: 70px; }
.drawn-ripples i:nth-child(3) { right: 52px; bottom: 61px; width: 94px; height: 50px; }

.captivity-copy h2 {
  font-size: clamp(2.8rem, 5.2vw, 4.8rem);
}

.captivity-copy > p:not(.eyebrow) {
  margin-bottom: 32px;
  color: #566d61;
}

.link-stack {
  border-top: 1px solid rgba(20, 40, 32, 0.22);
}

.link-stack a {
  display: flex;
  align-items: center;
  justify-content: space-between;
  min-height: 62px;
  border-bottom: 1px solid rgba(20, 40, 32, 0.22);
  font-size: 0.9rem;
  font-weight: 700;
}

.link-stack b {
  font-size: 1.12rem;
  font-weight: 500;
  transition: transform 180ms ease;
}

.link-stack a:hover b,
.link-stack a:focus-visible b {
  transform: translateX(5px);
}

.collaboration {
  background: var(--paper);
}

.collaboration-heading {
  align-items: end;
}

.profile-grid {
  display: grid;
  grid-template-columns: repeat(4, 1fr);
  border-top: 1px solid var(--line);
  border-left: 1px solid var(--line);
}

.profile-card {
  display: flex;
  flex-direction: column;
  min-height: 375px;
  padding: 30px;
  border-right: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
  transition: color 200ms ease, background-color 200ms ease;
}

.profile-card:hover,
.profile-card:focus-visible {
  color: white;
  background: var(--forest);
}

.profile-icon {
  display: grid;
  width: 54px;
  height: 54px;
  margin-bottom: 56px;
  place-items: center;
  color: var(--forest);
  background: var(--acid);
  border-radius: 42% 58% 51% 49% / 55% 38% 62% 45%;
  font-family: var(--serif);
  font-size: 1.35rem;
}

.profile-card:nth-child(2) .profile-icon { transform: rotate(8deg); }
.profile-card:nth-child(3) .profile-icon { transform: rotate(-7deg); }
.profile-card:nth-child(4) .profile-icon { transform: rotate(4deg); }

.profile-card h3 {
  margin-bottom: 12px;
  font-family: var(--serif);
  font-size: 1.5rem;
  font-weight: 500;
  letter-spacing: -0.03em;
}

.profile-card p {
  color: #677b71;
  font-size: 0.85rem;
}

.profile-card:hover p,
.profile-card:focus-visible p {
  color: rgba(255, 255, 255, 0.68);
}

.profile-link {
  margin-top: auto;
  font-size: 0.75rem;
  font-weight: 800;
}

.network {
  color: white;
  background: var(--forest);
}

.network-grid {
  display: grid;
  grid-template-columns: minmax(300px, 0.8fr) minmax(560px, 1.2fr);
  align-items: center;
  gap: 80px;
}

.network-copy h2 {
  max-width: 500px;
  font-size: clamp(3rem, 5.2vw, 5rem);
}

.network-copy > p:not(.eyebrow) {
  max-width: 500px;
  color: rgba(255, 255, 255, 0.68);
}

.metrics {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  margin: 0;
  border-top: 1px solid rgba(255, 255, 255, 0.18);
  border-left: 1px solid rgba(255, 255, 255, 0.18);
}

.metrics div {
  min-height: 190px;
  padding: 30px;
  border-right: 1px solid rgba(255, 255, 255, 0.18);
  border-bottom: 1px solid rgba(255, 255, 255, 0.18);
}

.metrics dt {
  margin-bottom: 9px;
  color: var(--acid);
  font-family: var(--serif);
  font-size: clamp(3.7rem, 6vw, 5.6rem);
  letter-spacing: -0.07em;
  line-height: 1;
}

.metrics dd {
  margin: 0;
  color: rgba(255, 255, 255, 0.68);
  font-size: 0.82rem;
  text-transform: lowercase;
}

.final-cta {
  position: relative;
  overflow: hidden;
  background: var(--acid);
}

.final-cta-inner {
  position: relative;
  z-index: 2;
  max-width: 920px;
  text-align: center;
}

.final-cta .eyebrow {
  color: rgba(20, 40, 32, 0.64);
}

.final-cta h2 {
  margin-bottom: 22px;
  font-size: clamp(3rem, 6.6vw, 6.4rem);
}

.final-cta p:not(.eyebrow) {
  max-width: 620px;
  margin: 0 auto 34px;
  color: rgba(20, 40, 32, 0.72);
}

.centered-buttons {
  justify-content: center;
}

.button-dark {
  color: white;
  background: var(--forest-deep);
  border-color: var(--forest-deep);
}

.button-dark:hover,
.button-dark:focus-visible {
  color: white;
  background: var(--forest);
  border-color: var(--forest);
}

.cta-sketch i {
  position: absolute;
  z-index: -1;
  display: block;
  border: 1px solid rgba(20, 40, 32, 0.16);
  border-radius: 50%;
}

.cta-sketch i:nth-child(1) { top: -110px; left: -240px; width: 520px; height: 520px; }
.cta-sketch i:nth-child(2) { top: -40px; left: -170px; width: 380px; height: 380px; }
.cta-sketch i:nth-child(3) { right: -250px; bottom: -250px; width: 580px; height: 580px; }
.cta-sketch i:nth-child(4) { right: -140px; bottom: -130px; width: 360px; height: 360px; }

.site-footer {
  padding: 76px 0 28px;
  color: white;
  background: #081c15;
}

.footer-grid {
  display: grid;
  grid-template-columns: 1.5fr 0.7fr 0.7fr 1fr;
  gap: 65px;
  padding-bottom: 70px;
}

.brand-footer {
  margin-bottom: 20px;
}

.footer-brand p,
.footer-contact p {
  max-width: 270px;
  color: rgba(255, 255, 255, 0.56);
  font-size: 0.82rem;
}

.footer-grid h2 {
  margin-bottom: 18px;
  color: rgba(255, 255, 255, 0.42);
  font-family: var(--sans);
  font-size: 0.67rem;
  font-weight: 800;
  letter-spacing: 0.16em;
  text-transform: uppercase;
}

.footer-grid > div:not(.footer-brand) > a {
  display: block;
  width: fit-content;
  margin-bottom: 8px;
  color: rgba(255, 255, 255, 0.8);
  font-size: 0.84rem;
}

.footer-grid a:hover,
.footer-grid a:focus-visible {
  color: var(--acid);
}

.footer-contact-link {
  padding-bottom: 4px;
  border-bottom: 1px solid rgba(255, 255, 255, 0.32);
  font-weight: 700;
}

.footer-bottom {
  display: flex;
  align-items: center;
  justify-content: space-between;
  padding-top: 26px;
  color: rgba(255, 255, 255, 0.35);
  border-top: 1px solid rgba(255, 255, 255, 0.12);
  font-size: 0.7rem;
}

:focus-visible {
  outline: 3px solid var(--acid);
  outline-offset: 4px;
}

@media (max-width: 1050px) {
  .desktop-nav {
    gap: 18px;
  }

  .disease-grid,
  .captivity-grid {
    gap: 55px;
  }

  .profile-grid {
    grid-template-columns: repeat(2, 1fr);
  }

  .network-grid {
    grid-template-columns: 1fr;
  }
}

@media (max-width: 860px) {
  .section {
    padding: 86px 0;
  }

  .desktop-nav,
  .header-inner > .button-small {
    display: none;
  }

  .mobile-menu {
    position: relative;
    display: block;
  }

  .mobile-menu summary {
    display: grid;
    width: 42px;
    height: 42px;
    place-content: center;
    gap: 5px;
    border: 1px solid rgba(255, 255, 255, 0.4);
    border-radius: 50%;
    cursor: pointer;
    list-style: none;
  }

  .mobile-menu summary::-webkit-details-marker { display: none; }

  .mobile-menu summary span {
    display: block;
    width: 17px;
    height: 1px;
    background: white;
  }

  .mobile-menu nav {
    position: absolute;
    top: 54px;
    right: 0;
    display: grid;
    width: min(300px, calc(100vw - 48px));
    padding: 18px;
    color: var(--ink);
    background: var(--paper);
    box-shadow: var(--shadow);
  }

  .mobile-menu nav a {
    padding: 11px 6px;
    border-bottom: 1px solid var(--line);
  }

  .hero {
    min-height: 820px;
  }

  .hero-copy {
    width: min(650px, 82%);
  }

  .hero-shade {
    background:
      linear-gradient(90deg, rgba(6, 25, 18, 0.95) 0%, rgba(6, 25, 18, 0.74) 54%, rgba(6, 25, 18, 0.14) 100%),
      linear-gradient(0deg, rgba(7, 25, 19, 0.58), transparent 48%);
  }

  .split-heading,
  .disease-grid,
  .captivity-grid {
    grid-template-columns: 1fr;
    gap: 40px;
  }

  .fact-grid,
  .action-grid {
    grid-template-columns: 1fr;
  }

  .fact-card {
    min-height: 335px;
    border-right: 0;
    border-bottom: 1px solid var(--line);
  }

  .fact-card:last-child {
    border-bottom: 0;
  }

  .action-card {
    min-height: 380px;
  }

  .disease-copy {
    max-width: 650px;
  }

  .disease-visual {
    min-height: 570px;
  }

  .captivity-grid {
    align-items: start;
  }

  .captivity-media {
    min-height: 520px;
  }

  .captivity-media > img {
    height: 500px;
  }

  .footer-grid {
    grid-template-columns: repeat(2, 1fr);
  }
}

@media (max-width: 620px) {
  .shell {
    width: min(100% - 34px, 1180px);
  }

  .section {
    padding: 70px 0;
  }

  .header-inner {
    min-height: 76px;
  }

  .brand {
    font-size: 1.15rem;
  }

  .brand-mark {
    width: 29px;
    height: 29px;
  }

  .hero {
    align-items: end;
    min-height: 810px;
  }

  .hero-image {
    object-position: 65% center;
  }

  .hero-shade {
    background: linear-gradient(0deg, rgba(5, 23, 16, 0.98) 5%, rgba(5, 23, 16, 0.82) 55%, rgba(5, 23, 16, 0.12) 100%);
  }

  .hero-inner {
    padding-top: 130px;
    padding-bottom: 140px;
  }

  .hero-copy {
    width: 100%;
  }

  .hero h1 {
    font-size: clamp(3.3rem, 15vw, 5rem);
  }

  .hero-note {
    left: 0;
    width: 100%;
    max-width: none;
    min-height: 72px;
    padding: 16px 18px;
  }

  .split-heading {
    gap: 24px;
  }

  .section-heading {
    margin-bottom: 42px;
  }

  .fact-card {
    padding: 30px 26px;
  }

  .disease-visual {
    min-height: 430px;
  }

  .photo-card-main {
    width: 88%;
    height: 320px;
  }

  .photo-card-small {
    width: 62%;
    height: 205px;
    border-width: 7px;
  }

  .orbit-one { width: 280px; height: 280px; }

  .action-card {
    min-height: 420px;
    padding: 28px;
  }

  .captivity-media {
    min-height: 430px;
  }

  .captivity-media > img {
    width: 100%;
    height: 410px;
  }

  .profile-grid,
  .metrics,
  .footer-grid {
    grid-template-columns: 1fr;
  }

  .profile-card {
    min-height: 310px;
  }

  .profile-icon {
    margin-bottom: 36px;
  }

  .network-grid {
    gap: 48px;
  }

  .metrics div {
    min-height: 155px;
  }

  .footer-grid {
    gap: 32px;
  }

  .footer-bottom {
    align-items: flex-start;
    flex-direction: column;
    gap: 8px;
  }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { transition-duration: 0.01ms !important; }
}